The Role of the Paleo-Critical Zone in Shaping Quaternary Hominin Evolution

The impact of changing environments on the evolution and dispersal of Homo sapiens is highly debated, but few data are available from equatorial Africa. Lake Victoria is the largest freshwater lake in the tropics and is currently a biogeographic barrier between the eastern and western branches of the East African Rift. The lake has previously desiccated at ~17 ka and again at ~15 ka, but little is known from this region prior to the Last Glacial Maximum. The Pleistocene terrestrial deposits on the northeast coast of Lake Victoria (94–36 ka) are ideal for paleoenvironmental reconstructions where volcaniclastic deposits (tuffs), fluvial deposits, tufa, and paleosols are exposed, which can be used to reconstruct Critical Zones (CZ) of the past (paleo-CZs). The paleo-CZ is a holistic concept that reconstructs the entire landscape using geologic records of the atmosphere, hydrosphere, lithosphere, biosphere, and pedosphere (the focus of this study). New paleosol-based mean annual precipitation (MAP) proxies from Karungu, Rusinga Island, and Mfangano Island indicate an average MAP of 750 ± 108 mm year⁻¹ (CALMAG), 800 ± 182 mm year⁻¹ (CIA-K), and 1,010 ± 228 mm year⁻¹ (PPM_1.0) with no statistical difference throughout the 11 m thick sequence. This corresponds to between 54 and 72% of modern precipitation. Tephras bracketing these paleosols have been correlated across seven sites, and sample a regional paleo-CZ across a ~55 km transect along the eastern shoreline of the modern lake. Given the sensitivity of Lake Victoria to precipitation, it is likely that the lake was significantly smaller than modern between 94 and 36 ka. This would have removed a major barrier for the movement of fauna (including early modern humans) and provided a dispersal corridor across the equator and between the rifts. It is also consistent with the associated fossil faunal assemblage indicative of semi-arid grasslands. During the Late Pleistocene, the combined geologic and paleontological evidence suggests a seasonally dry, open grassland environment for the Lake Victoria region that is significantly drier than today, which may have facilitated human and faunal dispersals across equatorial East Africa.

The abundance of the southern Pacific mollusk loco (Concholepas concholepas), among other conspicuous marine supplies, are often cited as critical resources behind the long-term cultural and demographic fluctuations of prehistoric hunter-gatherers in the coastal Atacama Desert. These societies inhabited one of the world's most productive marine environments flanked by one the world's driest deserts. Both of these environments have witnessed significant ecological variation since people first colonized them at the end of the Pleistocene (c. 13,000 cal yr BP). Here, we examine the relationship between the relative abundance of shellfish (a staple resource) along a 9,500-year sequence of archeological shell midden accumulations at Caleta (a small inlet or cove) Vitor, with past demographic trends (established via summed probability distributions of radiocarbon ages) and technological innovations together with paleoceanographic data on past primary productivity. We find that shellfish extraction varied considerably from one cultural period to the next in terms of the number of species and their abundance, with diversity increasing during periods of regionally decreased productivity. Such shifts in consumption patterns are considered community based management decisions, and for the most part they were synchronous with large and unusual regional demographic fluctuations experienced by prehistoric coastal societies in northern Chile. When taken together with their technological innovations, our data illustrates how these human groups tailored their socio-cultural patterns to what were often abrupt and prolonged environmental changes throughout the Holocene.