Dr. Hardingham received his degree in Biochemistry from the University of Cambridge. He went on to do a PhD with Professor Hilmar Bading at the MRC Laboratory of Molecular Biology, Cambridge on spatial aspects of Ca2+signalling in excitable cell lines. In 1998 he obtained an MRC Research Fellowship to continue working at the LMB on activity-dependent gene expression in neurons. He was also a Fellow of Clare College, Cambridge and Tutor in Molecular Biology. In 2002 he moved to the University of Edinburgh and obtained a Royal Society Research Fellowship to investigate aspects of activity-dependent gene expression and their impact on neuronal fate and function. He is currently a Reader in Neuroscience.